Humans in the Holocene
The Last Glacial Maximum
The Last Glacial Maximum (LGM) marks a pivotal period in Earth's history when ice sheets reached their greatest extent, approximately 26,500 to 19,000 years ago. During this time, vast ice sheets covered much of North America, Northern Europe, and other parts of the globe, dramatically altering the planet's climate and geography.

The climate of the LGM was significantly colder and drier than today. Temperatures were as much as 10 degrees Celsius cooler in some regions, fundamentally affecting global weather patterns and even the circulation of ocean currents. This led to a stark landscape, where much of today’s fertile land was barren tundra or covered by ice. The drop in temperatures and the expansion of ice sheets also caused a significant reduction in sea levels, revealing land bridges between continents, such as the famous Bering Land Bridge between Asia and North America. These bridges facilitated migrations of humans and animals, enabling them to move across continents.
The harsh conditions of the LGM forced flora and fauna to adapt, migrate, or face extinction. Many species migrated southward or to lower elevations to escape the encroaching ice, while others evolved to cope with the colder climate. This period of glaciation left a lasting impact on species distribution and evolution, evidence of which is observed in the genetic patterns of many modern species.
Humans living during the LGM also adapted to these severe climates. Archaeological evidence shows that human populations employed innovative survival strategies, from developing specialized hunting techniques to creating more efficient clothing and shelter. These adaptations were crucial for survival in a world where resources were scarce and the environment unforgiving.
As the climate eventually warmed, the ice sheets began to retreat, leading to the current interglacial period where we now live. The retreat of the ice transformed landscapes, created new habitats, and reshaped the courses of rivers and coastlines. This post-glacial world saw rising sea levels and a warmer climate, which played a key role in the development of modern ecosystems and human civilizations.
Understanding the Last Glacial Maximum is essential for scientists studying past climate changes and predicting future climate patterns. It provides a stark example of Earth's dynamic climate system and its profound impact on the biosphere and human societies.
Expansion into Northern Eurasia

Following the Last Glacial Maximum (LGM), which peaked around 20,000 years ago, the massive ice sheets that had covered much of northern Eurasia began to retreat. This marked the end of a cold phase in Earth's history, during which large portions of the continent had been uninhabitable to humans. As the ice receded, it exposed vast new territories that would become crucial for human habitation. The warming climate allowed for the gradual expansion of human populations into regions that had been covered by ice for millennia.
One of the key areas of human expansion was the region now known as Northern Europe, including the Scandinavian Peninsula. As the glaciers retreated, they left behind newly accessible coastlines, river valleys, and plains. The first groups to colonize these regions were hunter-gatherers, who followed migrating herds of large game such as reindeer and mammoths. These early settlers included the ancestors of modern populations like the Sami people, who adapted to the forested landscapes and utilized marine resources, especially in coastal areas. Archaeological evidence suggests that these early humans had sophisticated hunting, fishing, and tool-making technologies.
To the east, the vast steppe plains of what is now Russia were also exposed by the melting ice. This region was eventually populated by hunter-gatherers who took advantage of the rich biodiversity of the steppe, which included large mammals like bison and wild horses, as well as plant resources suitable for foraging. The rivers that traversed these plains, including the Volga, Don, and Dnieper, became vital routes for transportation, trade, and communication, facilitating the movement of goods and people across this expansive territory.
In Siberia, human populations began to enter the region as the ice melted and the climate warmed. These migrations occurred in phases, with some groups crossing the Bering Land Bridge from North America, while others moved eastward from Central Asia and Eastern Europe. The ancestors of the Inuit and other Arctic populations are believed to have descended from these early Siberian migrants. These peoples were well-adapted to the harsh, cold environments of Siberia and later the Arctic, developing technologies and subsistence strategies such as hunting, fishing, and herding that allowed them to survive in the region's extreme conditions.
The colonization of northern Eurasia after the Last Ice Age was not a singular event but rather a complex, gradual process involving multiple waves of migration, interaction, and adaptation. The interactions between different groups—whether newcomers or indigenous populations—led to a dynamic cultural exchange. Archaeological evidence shows a blending of tool-making technologies, artistic styles, and burial practices across the newly populated regions, reflecting a complex pattern of social and cultural exchanges.
The settlement of northern Eurasia after the retreat of the glaciers is a testament to human resilience and adaptability. It illustrates the profound impact that changing climates and landscapes can have on human history. As the ice sheets receded, they opened up new pathways for migration and settlement, setting the stage for millennia of cultural development and interaction across some of the most challenging and diverse environments on Earth.
The Colonization of the Americas

The colonization of the Americas by human populations is a vibrant field of ongoing research and debate. The prevailing theory is that the first inhabitants migrated from northeastern Asia via the Bering Land Bridge, a landmass exposed during the last Ice Age that connected Siberia to Alaska. This migration is believed to have occurred at least 15,000 years ago, but recent archaeological evidence suggests an even earlier human presence in North America.
One of the most compelling discoveries challenging the Clovis-first model is the fossilized footprints found at White Sands National Park in New Mexico, dated to between 20,000 and 23,000 years ago. These findings indicate that humans were present in the Americas long before the Clovis culture emerged, leading to a reexamination of the migration timelines and the routes early humans may have taken.
The migration patterns of these early populations were complex, involving multiple waves and varied pathways, including both coastal and inland routes. It is now increasingly recognized that people likely arrived in successive waves, with each group adapting to different environmental challenges related to the shifting climate of the Ice Ages. Some genetic and archaeological evidence suggests that the earliest wave of settlers may have made it all the way to the southernmost parts of South America, leaving behind a legacy in the genetic makeup of populations such as the Yámana, Kawésqar, and other Indigenous groups of Patagonia. This theory is supported by findings indicating that populations in the far south are genetically distinct from those in more northern regions, suggesting an earlier separation.
Advances in genetic research further illuminate the dynamic process of population expansion and diversification that led to the rich mosaic of Indigenous cultures present when Europeans arrived in the 15th and 16th centuries. The southern migration, in particular, demonstrates the remarkable adaptability of early humans who managed to reach some of the most remote and challenging environments in the world.
This narrative of human settlement underscores the adaptability and resilience of early human populations, marking one of the most significant migration and colonization events in human history. Indigenous oral traditions, which often recount ancient migrations and ancestral origins, provide invaluable insights that complement archaeological and genetic evidence, enriching our understanding of this pivotal chapter in human history.
The Green Sahara: A Haven for Early Human Settlement and Innovation
The pathogenic environment closer to the equator has always posed significant challenges for human populations. High humidity, abundant water sources, and dense vegetation make equatorial regions ideal breeding grounds for various disease-carrying organisms, such as mosquitoes. As a result, the burden of diseases like malaria, sleeping sickness, and other infections has historically been much higher in these areas. The African Humid Period, roughly between 14,600 and 5,000 years ago, was a time when the Sahara Desert was lush, green, and teeming with life. The region experienced a dramatically different climate, characterized by ample rainfall, extensive lakes, and flourishing grasslands. In this fertile environment, plants and animals thrived, and it is plausible that human communities settled and began to exploit the newfound abundance. The drier savannas and grasslands of the Sahara would have had a significantly reduced pathogenic load compared to the equatorial regions, potentially offering a healthier environment for human settlement and development. This freedom from high disease pressure could have been a key factor that allowed communities to flourish, invest time in cultivating crops, raising animals, and building more permanent settlements.

The African Humid Period coincided with the emergence of the first villages and the advent of agriculture in various parts of the Mediterranean, leading many researchers to speculate that similar developments might have taken place within the Sahara as well. While archaeological evidence from the Sahara is more challenging to uncover due to the subsequent return of arid conditions and the covering sands, there are indications that people lived in the region in ways similar to early agriculturalists elsewhere. The humid landscape of the Sahara likely supported communities engaged in early forms of plant cultivation and livestock herding, much like those found in other parts of North Africa and the Near East during the same time frame. The presence of tools, pottery, and rock art depicting cattle hints at the existence of societies that may have practiced a form of agriculture or animal husbandry, taking advantage of the temporary bounty of the region. These communities enjoyed the benefits of a fertile landscape, offering a respite from the harsher conditions and higher pathogenic load found farther south in tropical Africa.
Pearl millet (Pennisetum glaucum), one of Africa’s oldest domesticated crops, provides a compelling example of this early agricultural development. It is believed to have been first domesticated in the southern regions of the Sahara or Sahel, where the conditions during the African Humid Period would have supported its cultivation. As the Sahara transitioned to a desert, pearl millet, with its remarkable drought tolerance, may have been adapted by these early communities to suit increasingly arid conditions. This crop’s domestication likely reflects the adaptation strategies of Saharan inhabitants who initially took advantage of the fertile and abundant landscape but later responded to environmental shifts by cultivating plants suited to harsher climates. Today, pearl millet remains a vital staple in arid regions, its history a possible legacy of the early agricultural experiments conducted within the Green Sahara.
The African Humid Period, therefore, presents a compelling scenario in which the Sahara served as a cradle for human innovation and adaptation. As the environment changed, humans adapted by forming settlements and possibly practicing early agriculture, just as their contemporaries were doing in other regions around the Mediterranean. The return of arid conditions eventually forced these communities to either adapt once again or migrate, but the legacy of this green Sahara era continues to shape our understanding of early human societies and their ability to thrive under dynamic environmental conditions.